Page:Descent of Man 1875.djvu/255

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Chap. VIII.
Sexual Selection.
239

do the sexes of the aboriginal parent-species; yet they acquire their characteristic plumage late in life, for the chickens are distinctly pencilled. With respect to other characters beside colour, in the wild-parent species and in most of the domestic breeds, the males alone possess a well-developed comb; but in the young of the Spanish fowl it is largely developed at a very early age, and, in accordance with this early development in the male, it is of unusual size in the adult female. In the Game breeds pugnacity is developed at a wonderfully early age, of which curious proofs could be given; and this character is transmitted to both sexes, so that the hens, from their extreme pugnacity, are now generally exhibited in separate pens. With the Polish breeds the bony protuberance of the skull which supports the crest is partially developed even before the chickens are hatched, and the crest itself soon begins to grow, though at first feebly;[1] and in this breed the adults of both sexes are characterised by a great bony protuberance and an immense crest.


Finally, from what we have now seen of the relation which exists in many natural species and domesticated races, between the period of the development of their characters and the manner of their transmission—for example, the striking fact of the early growth of the horns in the reindeer, in which both sexes bear horns, in comparison with their much later growth in the other species in which the male alone bears horns—we may conclude that one, though not the sole cause of characters being exclusively inherited by one sex, is their development at a late age. And secondly, that one, though apparently a less efficient cause of characters being inherited by both sexes, is their development at an early age, whilst the sexes differ but little in constitution. It appears, however, that some difference must exist between the sexes even during a very early embryonic period, for characters developed at this age not rarely become attached to one sex.


Summary and concluding remarks.—From the foregoing discussion on the various laws of inheritance, we learn that the characters of the parents often, or even generally, tend to become developed in the offspring of the same sex, at the same age, and periodically at the same season of the year, in which they first

  1. For full particulars and references on all these points respecting the several breeds of the Fowl, see 'Variation of Animals and Plants under Domestication,' vol. i. pp. 250, 256. In regard to the higher animals, the sexual differences which have arisen under domestication are described in the same work under the head of each species.