which is opposite to the female, and which would otherwise be concealed, so that nearly all the beautifully spotted feathers are exhibited at the same time.
The Argus pheasant affords a much more remarkable case. The immensely developed secondary wing-feathers are confined to the male; and each is ornamented with a row of from twenty to twenty-three ocelli, above an inch in diameter. These feathers are also elegantly marked with oblique stripes and rows of spots of a dark colour, like those on the skin of a tiger and leopard combined. These beautiful ornaments are hidden until the male shews himself off before the female. He then erects his tail, and expands his wing-feathers into a great, almost upright, circular fan or shield, which is carried in front of the body. The neck and head are held on one side, so that they are concealed by the fan; but the bird in order to see the female, before whom he is displaying himself, sometimes pushes his head between two of the long wing-feathers (as Mr. Bartlett has seen), and then presents a grotesque appearance. This must be a frequent habit with the bird in a state of nature, for Mr. Bartlett and his son on examining some perfect skins sent from the East, found a place between two of the feathers, which was much frayed, as if the head had here frequently been pushed through. Mr. Wood thinks that the male can also peep at the female on one side, beyond the margin of the fan.
The ocelli on the wing-feathers are wonderful objects; for they are so shaded that, as the Duke of Argyll remarks,[1] they stand out like balls lying loosely within sockets. When I looked at the specimen in the British Museum, which is mounted with the wings expanded and trailing downwards, I was however greatly disappointed, for the ocelli appeared flat, or even concave. But Mr. Gould soon made the case clear to me, for he held the feathers erect, in the position in which they would naturally be displayed, and now, from the light shining on them from above, each ocellus at once resembled the ornament called a ball and socket. These feathers have been shewn to several artists, and all have expressed their admiration at the perfect shading. It may well be asked, could such artistically shaded ornaments have been formed by means of sexual selection? But it will be convenient to defer giving an answer to this question, until we treat in the next chapter of the principle of gradation.
The foregoing remarks relate to the secondary wing-feathers, but the primary wing-feathers, which in most gallinaceous birds are uniformly coloured, are in the Argus pheasant equally wonderful. They are of a soft brown tint with numerous dark
- ↑ 'The Reign of Law,' 1867, p. 203.
