be better able than the mediocres to crush through the thick grass and foliage whenever he was scampering for his life, to jump over obstacles, and possibly to run somewhat faster than they. So far as he is small of his kind, he would be better able to run through narrow openings, to make quick turns, and to hide himself. Under the general circumstances it would be found that animals of some particular stature had on the whole a better chance of escape than any other; and if their race is closely adapted to these circumstances in respect to stature, the most favored stature would be identical with the mean of the race. Though the impediments to flight are less unfavorable to this (stature) than to any other, they will differ in different experiences. The course of an animal might chance to pass through denser foliage than usual, or the obstacles in his way may be higher. In that case an animal whose stature exceeded the mean would have an advantage over mediocrities. Conversely the circumstances might be more favorable to a small animal. Each particular line of escape might be most favorable to some particular stature, and, whatever this might be, it might in some cases be more favored than any other. But the accidents of foliage and soil in a country are characteristic and persistent, and may fairly be considered as approximating to a typical kind. Therefore those which most favor the animals of the mean stature will be more frequently met with than those which favor any other stature, and the frequency of the latter occurrence will diminish rapidly as the stature departs from the mean.
"It might well be that natural selection would favor the indefinite increase of numerous separate faculties if their improvement could be effected without detriment to the rest: then mediocrity in that faculty would not be the safest condition. Thus an increase of fleetness would be a clear gain to an animal liable to be hunted by beasts of prey, if no other useful faculty v/as thereby diminished.
"But a too free use of this 'if' would show a jaunty disregard of a real difficulty. Organisms are so knit together that change in one direction involves change in many others; these may not attract attention, but they are none the less existent. Organisms are like ships of war, constructed for a particular purpose in warfare as cruisers, line-of-battle-ships, etc., on the principle of obtaining the utmost efficiency for their special purpose. The result is a compromise between a variety of conflicting desiderata, such as cost, speed, accommodation, stability, weight of guns, thickness of armor, quick steering power, and so on. It is hardly possible in a ship of any established type to make an improvement in any one of these respects without a sacrifice in other directions. If the fleetness is increased, the engines must be
