Page:Darwinism by Alfred Wallace 1889.djvu/148

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CHAPTER VI

DIFFICULTIES AND OBJECTIONS

Difficulty as to smallness of variations—As to the right variations occurring when required—The beginnings of important organs—The mammary glands—The eyes of flatfish—Origin of the eye—Useless or non-adaptive characters—Recent extension of the region of utility in plants—The same in animals—Uses of tails—Of the horns of deer—Of the scale-ornamentation of reptiles—Instability of non-adaptive characters—Delbœuf's law—No "specific" character proved to be useless—The swamping effects of intercrossing—Isolation as preventing intercrossing—Gulick on the effects of isolation—Cases in which isolation is ineffective.

In the present chapter I propose to discuss the more obvious and often repeated objections to Darwin's theory, and to show how far they affect its character as a true and sufficient explanation of the origin of species. The more recondite difficulties, affecting such fundamental questions as the causes and laws of variability, will be left for a future chapter, after we have become better acquainted with the applications of the theory to the more important adaptations and correlations of animal and plant life.

One of the earliest and most often repeated objections was, that it was difficult "to imagine a reason why variations tending in an infinitesimal degree in any special direction should be preserved," or to believe that the complex adaptation of living organisms could have been produced "by infinitesimal beginnings." Now this term "infinitesimal," used by a well-known early critic of the Origin of Species, was never made use of by Darwin himself, who spoke only of variations being "slight," and of the "small amount" of the variations that might be selected. Even in using these terms he undoubtedly afforded