Page:History of botany (Sachs; Garnsey).djvu/532

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512
Theory of the Nutrition
[BOOK III.


up together, the weeping of severed root-stocks and the rise of the sap in the wood in transpiring plants. The first is caused, he thinks, by impulsion, the other by attraction; we should now say, that in weeping root-stocks the water is pressed upwards, in transpiring plants drawn up. He then refers the phenomenon of impulsion to endosmose in the roots, and without going much into detail as regards the anatomical conditions, he compares a weeping root-stock to his own endosmometer, in the tube of which the fluid that has been sucked in rises by endosmose and even flows over; it is true that no very thorough understanding of the matter was gained in this way, but at any rate the principle which was to explain it was indicated. He then endeavours to explain the movement of the water which ascends in the wood of transpiring plants by the action of endosmose from cell to cell. In this he failed entirely, as was afterwards perceived; but he succeeded in showing that all the mechanical explanations that had been previously attempted were incorrect, and the whole treatise, though unsatisfactory in its main result, contains a great number of ingenious experiments and acute remarks.

With the exception of Théodore de Saussure, who occupied himself exclusively with chemical questions in physiology, Dutrochet was the only vegetable physiologist in the period between 1820 and 1840 who studied all its more important questions thoroughly and experimentally; his treatise on the respiration of plants, which has been already mentioned, is excellent in itself, and was of the greatest importance at the time it appeared, because it brought the chemical processes in respiration, the entrance and exit of the gases, for the first time into correct connection with the air-passages in the plant, with the stomata, the vessels, and the intercellular spaces, and submitted the composition of the air contained in the cavities of plants to careful examination. It was the best work on the respiration of plants in the year 1837 and for a long time after; and if Dutrochet made the mistake of regarding the oxygen