The formation of vegetable mould, through the action of worms

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THE FORMATION

OF

VEGETABLE MOULD,

THROUGH THE

ACTION OF WORMS,

WITH

OBSERVATIONS ON THEIR HABITS.


By CHARLES DARWIN, LL. D., F.R.S.


WITH ILLUSTRATIONS


LONDON:
JOHN MURRAY, ALBEMARLE STREET.
1881.

The right of Translation is reserved.

LONDON:

PRINTED BY WILLIAM CLOWES AND SONS, LIMITED,

STANFORD STREET AND CHARING CROSS.

CONTENTS.

INTRODUCTION
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Page 1–7

CHAPTER I.

HABITS OF WORMS.

Nature of the sites inhabited—Can live long under water—Nocturnal—Wander about at night—Often lie close to the mouths of their burrows, and are thus destroyed in large numbers by birds—Structure—Do not possess eyes, but can distinguish between light and darkness—Retreat rapidly when brightly illuminated, not by a reflex action—Power of attention—Sensitive to heat and cold—Completely deaf—Sensitive to vibrations and to touch—Feeble power of smell—Taste—Mental qualities—Nature of food—Omnivorous—Digestion—Leaves before being swallowed, moistened with a fluid of the nature of the pancreatic secretion—Extra-stomachal digestion—Calciferous glands, structure of—Calcareous concretions formed in the anterior pair of glands—The calcareous matter primarily an excretion, but secondarily serves to neutralise the acids generated during the digestive process 8–54

CHAPTER II.

HABITS OF WORMS.continued.

Manner in which worms seize objects—Their power of suction—The instinct of plugging up the mouths of their burrows—Stones piled over the burrows—The advantages thus gained—Intelligence shown by worms in their manner of plugging up their burrows—Various kinds of leaves and other objects thus used—Triangles of paper—Summary of reasons for believing that worms exhibit some intelligence—Means by which they excavate their burrows, by pushing away the earth and swallowing it—Earth also swallowed for the nutritious matter which it contains—Depth to which worms burrow, and the construction of their burrows—Burrows lined with castings, and in the upper part with leaves—The lowest part paved with little stones or seeds—Manner in which the castings are ejected—The collapse of old burrows—Distribution of worms—Tower-like castings in Bengal—Gigantic castings on the Kilgiri Mountains—Castings ejected in all countries 55–128

CHAPTER III.

THE AMOUNT OF FINE EARTH BROUGHT UP BY WORMS TO THE SURFACE.

Rate at which various objects strewed on the surface of grass-fields are covered up by the castings of worms—The burial of a paved path—The slow subsidence of great stones left on the surface—The number of worms which live within a given space—The weight of earth ejected from a burrow, and from all the burrows within a given space—The thickness of the layer of mould which the castings on a given space would form within a given time if uniformly spread out—The slow rate at which mould can increase to a great thickness—Conclusion 129–175

CHAPTER IV.

THE PART WHICH WORMS HAVE PLAYED IN THE BURIAL OF ANCIENT BUILDINGS.

The accumulation of rubbish on the sites of great cities independent of the action of worms—The burial of a Roman villa at Abinger—The floors and walls penetrated by worms—Subsidence of a modern pavement—The buried pavement at Beaulieu Abbey—Roman villas at Chedworth and Brading—The remains of the Roman town at Silchester—The nature of the débris by which the remains are covered—The penetration of the tesselated floors and walls by worms—Subsidence of the floors—Thickness of the mould—The old Roman city of Wroxeter—Thickness of the mould—Depth of the foundations of some of the buildings—Conclusion 176–229

CHAPTER V.

THE ACTION OF WORMS IN THE DENUDATION OF THE LAND.

Evidence of the amount of denudation which the land has undergone—Subaerial denudation—The deposition of dust—Vegetable mould, its dark colour and fine texture largely due to the action of worms—The disintegration of rocks by the humus-acids—Similar acids apparently generated within the bodies of worms—The action of these acids facilitated by the continued movement of the particles of earth—A thick bed of mould checks the disintegration of the underlying soil and rocks—Particles of stone worn or triturated in the gizzards of worms—Swallowed stones serve as millstones—The levigated state of the castings—Fragments of brick in the castings over ancient buildings well rounded. The triturating power of worms not quite insignificant under a geological point of view 230–258

CHAPTER VI.

THE DENUDATION OF THE LANDcontinued.

Denudation aided by recently ejected castings flowing down inclined grass-covered surfaces—The amount of earth which annually flows downwards—The effect of tropical rain on worm castings—The finest particles of earth washed completely away from castings—The disintegration of dried castings into pellets, and their rolling down inclined surfaces—The formation of little ledges on hill-sides, in part due to the accumulation of disintegrated castings—Castings blown to leeward over level land—An attempt to estimate the amount thus blown—The degradation of ancient encampments and tumuli—The preservation of the crowns and furrows on land anciently ploughed—The formation and amount of mould over the Chalk formation 259—304

CHAPTER VII.

CONCLUSION.

Summary of the part which worms have played in the history of the world—Their aid in the disintegration of rocks—In the denudation of the land—In the preservation of ancient remains—In the preparation of the soil for the growth of plants—Mental powers of worms—Conclusion 305–313

Index
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315–326

This work was published before January 1, 1929, and is in the public domain worldwide because the author died at least 100 years ago.

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