explained by reference to the traces yielded by seismometers. Perhaps the greatest triumph in seismological investigation has been the determination of the varying rates at which motion is propagated through the world. These measurements have already thrown new light upon its effective rigidity, and if we assume that the density of the earth increases uniformly from its surface towards its centre, so that its mean density is 5·5, then, according to Knott, the coefficient of elasticity which governs the transmission of preliminary tremors of an earthquake increases at a rate of nearly 1·2% per mile of descent. (J. Mi.)
Authorities.—J. Milne, Seismology (London, 1898), Earthquakes (London, 1898), Bakerian Lecture, “Recent Advances in Seismology,” Proc. Roy. Soc., 1906, 77, p. 365; J. A. Ewing, Memoir on Earthquake Measurement (Tokyo, 1883); C. E. Dutton, Earthquakes in the Light of the New Seismology (London, 1904); “The Charleston Earthquake of Aug. 31, 1886,” Ninth Annual Report of the United States Geological Survey, 1889; W. H. Hobbs, Earthquakes, an Introduction to Seismic Geology (London, 1908), “The San Francisco Earthquake and Fire, 1906,” Bull. U.S. Geol. Surv. No. 324; “The California Earthquake of Ap. 18, 1906,” Rep. State Earthq. Com. (Washington, D.C., 1908); R. D. Oldham, “Report on the Great Earthquake of 12 June 1897,” Mem. Geol. Surv. India, xxix. 1899, “On the Propagation of Earthquake Motion to great Distances,” Phil. Trans., 1900, A, vol. 194, p. 135, “The Constitution of the Interior of the Earth as revealed by Earthquakes,” Quar. Jour. Geol. Soc., 1906, 62, p. 456; 1907, 63, p. 344; C. Davison, A Study of Recent Earthquakes (London, 1905); The Hereford Earthquake of December 17, 1896 (Birmingham, 1899), “The Investigation of Earthquakes,” Beiträge z. Geophysik, Bd. ix., 1908, p. 201, and papers on British earthquakes in Quart. Jour. Geol. Soc.; T. J. J. See, “The Cause of Earthquakes, Mountain Formation and Kindred Phenomena connected with the Physics of the Earth,” Proc. Amer. Phil. Soc., 1906, 45, p. 273; F. Frech, “Erdbeben und Gebirgsbau,” Petermann’s Mitteilungen, Bd. 53, 1907, p. 245 (with maps); C. G. Knott, The Physics of Earthquake Phenomena (Oxford, 1908); Comte F. de Montessus de Ballore, Les Tremblements de terre: géographie séismologique (Paris, 1906), La Science séismologique (1907); Transactions of the Seismological Society of Japan; Seismological Journal (Yokohama); Bollettino della Società Sismologica Italiana (Rome); Reports of the British Association, containing the annual reports of the Committee for Seismological Investigations; papers in the Beiträge zur Geophysik and the Ergänzungsbände.
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| From Strasburger’s Lehrbuch der Botanik, by permission of Gustav Fischer. |
| Geaster Granulosus, nat. size. |
EARTH-STAR (Geaster), in botany, a kind of puff-ball, with a distinct outer coat which, on separating from the inner, splits into several divisions, which become reflexed and spread like a star. The inner coat enveloping the spores is supported, like a ball, either with or without a stalk on the upper face of the star. The spores escape generally by means of a distinct aperture which appears in the top of the ball. There are several species in Britain found on the ground or on decaying leaves. They are rare or local, but more common in the south or south-east of England than in other parts of Britain.
EARTHWORM, the common name of a chaetopod worm found nearly all over the world. Linnaeus recognized only one species of earthworm and named it Lumbricus terrestris. There are now one thousand well-characterized species known from different parts of the world, and the number increases almost daily. The earthworms of England belong entirely to the three genera Lumbricus, Allolobophora and Allurus, which are further subdivided by some systematists; and these genera form the prevalent earthworm fauna of the Palaearctic region and are also very numerous in the Nearctic region. Elsewhere they do not appear to be indigenous, but are replaced by the numerous other genera of the families enumerated in the article Chaetopoda (q.v.). It is a remarkable fact that these genera, comprizing a separate family Lumbricidae, when introduced into tropical and other countries, thrive abundantly and oust the indigenous forms. In gatherings of earthworms from various extra-European countries it is always found that if the collections have been made in cultivated ground and near the coast the worms are of European species; farther inland the native forms are met with. Inasmuch as in every case the Lumbricidae from non-European countries are identical with European species, since it has been shown that these animals are very readily introduced accidentally with plants, &c., and in view of the fact that they are impatient of sea water, it seems clear that the presence of these Lumbricidae in other continents is due to accidental transportation. Most earthworms live in the soil, which they devour as they burrow through it. A few, like their allies the river worms (Limicolae), habitually frequent streams, lakes, &c. One genus, at any rate, viz. Pontodrilus, seeks an unusual environment, and is found in heaps of sea-weed cast up by the sea. The range of this genus is therefore naturally wider than that of other genera which are confined to land masses and cannot cross the sea by their own efforts. It might be inferred, therefore, and the inference is proved by facts, that truly oceanic islands have no indigenous fauna of earthworms, but are inhabited by forms which are identical with those of neighbouring continents, and doubtless, therefore, accidentally introduced.
Like the leeches the earthworms produce cocoons which are a product of the glandular epithelium of the clitellum. In these cocoons are deposited the eggs together with a certain amount of albumen upon which the developing embryos feed. So far as is known, the production of cocoons is universal among earthworms and the remaining Oligochaeta of aquatic habit. The young leave the cocoon as fully formed earthworms in which, however, the genitalia are not fully developed. There is no free living larval stage. Out of a single cocoon emerge a varying number of young worms, the numbers being apparently characteristic of the species. The work of earthworms in aiding in the production of the subsoil and in levelling the surface was first studied by C. Darwin, and has since been investigated by others. This work is partly carried out beneath the surface and partly on the surface, upon which the worms wander at night and eject the swallowed and triturated earth; frequently castings of some height are formed of coiled ropes of agglutinated particles of mould. The indigenous species of Great Britain, about twenty in number, do not grow to a greater length than some 10 in.; but in several tropical countries there are species which grow to a length of from 3 to 6 ft. Thus we have in Natal the gigantic Microchaeta rappi, in Ceylon Megascolex coeruleus, in Australia Megascolides australis, and an equally large form in South America. (F. E. B.)
EARWIG, an insect belonging to the Forficulidae, a family usually referred to the Orthoptera, but sometimes regarded as typifying a special order, to which the names Dermaptera, Dermatoptera and Euplexoptera have been given, in allusion to certain peculiarities in the structure of the wings in the species that possess them. The front wings are short and horny and when at rest meet without overlapping in the middle line, like the wing-cases of brachelytrous (cocktail) beetles. The hind wings, on the contrary, are for the most part membranous and, when extended, of large size; each consists of two portions, the distal of which, in virtue of the arrangement and jointing of its nervures, is capable of being both doubled up and folded fanwise beneath the proximal, which is partly horny when the wing is tucked away under the front wing-case of the same side. Apart from these characteristics, the most distinctive feature of earwigs is the presence at the end of the abdomen of a pair of pincers which are in reality modified appendages, known as cercopods, and represent the similar limbs of Japyx and the caudal feelers of Campodea and some other insects.
The Forficulidae are almost cosmopolitan; but the various species and genera differ from each other both in structure and size to a comparatively slight extent. The length and armature of the pincers and the presence or absence of wings are perhaps the most important features used by systematists in distinguishing the various kinds. Of particular zoological interest in this connexion is a Ceylonese genus Dyscritina, in which the cercopods are long, many-jointed and filiform during the early stages of growth, and only assume at the last moult the forcipate structure characteristic of the family. The best known earwig is the common European species, Forficula auricularia. This insect is gregarious and nocturnal. It hides by day under stones or
