Page:Life and death (1911).djvu/126

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  • tion or simplification of the substance of reserve takes

place. Here is something that meets the case, and we may note the coincidence. It does not mean that the disposable energy is really used to increase the protoplasm, nor that the protoplasm itself is thereby increased. It merely signifies that the wherewithal exists to provide for that increase if it takes place.

It is therefore possible that the active protoplasm follows the law of functional assimilation; but it is certain that the reserve-stuff follows the law laid down by Claude Bernard. All these considerations definitely result in the confirmation of this second law of general physiology, according to which all vital energies are borrowed from the potential chemical energy of the reserve-stuff of alimentary origin. § 4. The Third Law of Biological Energetics. The third law of biological energetics is also drawn from experiment. It relates no longer to the point of departure of the cycle of animal energy, but to its final position. The energetic transformations of the animal end in thermal energy.

This is the most novel part of the theory, and, if we may say so, that least understood by physiologists themselves. The energy resulting from the chemical potential of food, having passed through the organism (or simply through the organ which we are considering in action), and having given rise to phenomenal appearances more or less diversified, more or less dim or clear, obscure or obvious, which are the characteristic or still irreducible manifestations of vitality, finally