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Pteridophytes had been gained, although the prothallia had been observed preceding the growth of the plant in Equisetum and Ferns. It was not till some years after Griffith's death that fuller facts as to the sexual organs were obtained and led to the right comparisons.
Griffith's work on the Bryophyta shows the same power of observation as that on the ovule, but the difficulties due to imperfect instruments are more evident. His views on reproduction were here, however, clear, since the development of the capsule was definitely related to the fertilisation of the pistilla (archegonia) by the substance formed in the anthers. His figures indicate how much he saw, and how here also he sought in development the interpretation of mature structure.
His early interest in the Liverworts, especially the Marchantiaceae, continued, and all the forms he collected were carefully examined and figured with his usual accuracy.
One of the Liverworts Griffith described may be taken as an illustration to this part of our subject on account of the interest of its re-discovery and re-description in 1910 by Goebel. This is a plant collected in Assam and named Monosolenium tenerum. This Marchantiaceous plant is described as having no air-chamber layer, as bearing sessile, dorsal, antheridial receptacles, and terminal, shortly stalked archegoniophores with one ventral groove in the stalk. A single archegonium—later capsule—is found in each of the half-dozen involucres. Spores and irregular bodies were found in the capsule.
Recently Goebel had two tea-plants sent home from Canton. They died, but he kept the soil moist on the chance of germinating seeds. Among a number of other plants there turned up a new Liverwort. On examination this proved to be Griffith's Monosolenium—all types of which had been lost—a most interesting form related to the Corsiniaceae.
In the Mosses and the Liverworts generally Griffith was clear on the development of the capsule or fruit following on the impregnation of an archegonium. But in Anthoceros, while he recognised the antheridia he was not clear as to the sunken archegonia, and regarded the capsules as arising by impregnation of unrecognisable spots on the young frond or thallus. He