common integument as in the Macropodidae. The hallux is opposable and nailless. The tail is nearly always long and prehensile. The pouch is well developed; the stomach not sacculated; a caecum is present (except in Tarsipes). These are really the principal distinctions between the two families. In addition, it may be mentioned that the lower incisors have not a scissor-like action as in the Kangaroos.
The Phalangers may be divided into four sub-families.
The first of these, that of the Phalangerinae, contains the genera Phalanger (including Cuscus), Acrobates, Distaechurus, Dromicia, Gymnobelideus, Petaurus, Petauroides, Dactylopsila, Pseudochirus, and Trichosurus.
These genera agree in the following generalities:—Tail well developed, often very long; three incisors above, and at least two premolars both above and below; caecum long and simple; stomach without a cardiac gland; liver not very complicated by secondary furrows, with a distinct caudate lobe; the vaginal median culs-de-sac often coalesced; lungs with an azygos lobe.
Fig. 68.—Bones of leg and foot of Phalanger. ast, Astragalus; calc, calcaneum; cub, cuboid; ect.cun, ecto-cuneiform; ent.cun, ento-cuneiform; fb, fibula; mes.cun, meso-cuneiform; nav, navicular; tib, tibia; I-V, first to fifth toes. (After Owen.)
The second sub-family, Phascolarctinae (with the Koala only), is thus characterised:—Tail rudimentary; cheek-pouches present; superior incisors three, but only one premolar above and below;
