Page:Miscellaneousbot01brow.djvu/465

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OF THE UNIMPREGNATED OVULUM.
447

coats of the ovulum before impregnation;[1] and of M. Turpin, as to the situation of the micropyle, and its being the cicatrix of a vascular cord. Yet he seems not to admit the function ascribed to it, and asserts that it is in many cases wanting.[2]


The account which I have given of the structure of the vegetable ovulum differs essentially from all those now quoted, and I am not acquainted with any other observations of importance respecting it.

Of the authors referred to, it may be remarked, that those who have most particularly attended to the ovulum externally, have not always examined it at a sufficiently [548 early period, and have confined themselves to its surface: that those who have most minutely examined its internal structure, have trusted too much to sections merely, and have neglected its appearance externally: and that those who have not at all examined it in the early stage have given the most correct account of its surface. This account was founded on a very limited observation of the seeds, generalized and extended to the unimpregnated ovulum, in connexion with an hypothesis then very commonly received: but this hypothesis being soon after abandoned, their statement respecting the ovulum was rejected along with it.

In the ovulum of Kingia, the inner membrane, with relation to the external umbilicus, is inverted; and this, as I have already observed, though in direct opposition to M. Turpin's account, is the usual structure of the organ. There are, however, several families in each of the two primary divisions of phænogamous plants, in which the inner membrane, and consequently the nucleus, agrees in direction with the testa. In such cases the external umbilicus alone affords a certain indication of the position of the future embryo.

It is an obvious consequence of what has been already stated, that the radicle of the embryo can never point directly to the external umbilicus or hilum, though this is

  1. Elém. Philos. Bot. p. 338.
  2. Id. p. 340.