Page:North American Plesiosaurs- Elasmosaurus, Cimoliasaurus, and Polycotylus.pdf/13

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S. W. Williston—North American Plesiosaurs.
233

Museum, obtained a good many years ago from the Fort Hays limestone, or basal strata of the Niobrara Cretaceous, of Jewell county, Kansas, by B. F. Mudge. The specimen bears the catalogue number 1640. Unfortunately, the specimen had been injured in collecting before it fell into proper hands. Originally it is probable that the larger part of the pectoral girdle, and perhaps, also, of the pelvic girdle and hind limb had been present, in addition to numerous vertebræ, and all in an undistorted condition. The specimen, notwithstanding what it has suffered, is of much interest, since it is the only vertebrate of which I have any knowledge from the Hays limestone. Additional figures and descriptions will be given later. For the present, the figures of the femur, ilia, and dorsal and sacral vertebræ given in Plate IV will render the species recognizable. A massive fragment of the scapula shows a broad and firm union with its mate in the middle line. The posterior projection of the coracoid is very long and much constricted before its extremity, its distal width being a little less than twice that of its least width; the outer posterior angle is acute and not much produced. The femur shows facets for but two epipodial bones.

Length of femur....................................................................................................................................................................................................................................................337mm

Greatest width distally....................................................................................................................................................................................................................................................206

A rather common species referable to this genus from the fence-post and lower horizons of the Benton is represented by a number of specimens in the Kansas University collections, and will be described later, with figures.

Two additional species also referable to this genus are known to me from the Hailey shales (probably equivalent to the Benton Cretaceous) of Wyoming, and will be described and figured later.

Polycotylus Cope.

The genus Polycotylus, described by Cope in 1870 from a number of mutilated vertebræ and fragments of podial bones, has remained hitherto much of a problem, and its characters have been very generally misunderstood. Fortunately, there is an excellent specimen in the Yale Museum (No. 1125), collected now many years ago by the late Professor Marsh in the vicinity of Fort Wallace, Kansas, from the Niobrara chalk, which I believe can be referred with certainty to the type species P. latipinnis Cope. That it belongs in the genus Polycotylus is beyond dispute, the vertebræ agreeing quite with the type as they do. This species seems to be the most common one of the order in the Kansas chalk, and is represented by several other specimens in the Yale Museum and by specimens in the University of Kansas collection. It is not at