Page:The Annals and Magazine of Natural History, vol. 16ss (1855).djvu/222

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190
Mr. A.R. Wallace on the Law which has regulated

furnish a countless number of analogous examples;—the Goliathi of Africa; the Ornithopteræ of the Indian islands, the Heliconide of South America, the Danaidæ of the East, and in all, the most closely allied species found in geographical proximity. The question forces itself upon every thinking mind,—why are these things so? They could not be as they are, had no law, regulated their creation and dispersion. The law here enunciated not merely explains, but necessitates the facts we see to exist, while the vast and long-continued geological changes of the earth readily account for the exceptions, and apparent diserepancies that here and there occur. The writer’s object in putting forward his views in the present imperfect manner is to submit them to the test of other minds, and to be made aware of all the facts supposed to be inconsistent with them. As his hypothesis is one which claims acceptance solely as explaining and connecting facts which exist in nature, he expects facts alone to be brought to disprove it; not a à-priori arguments against its probabiliy.

The phænomena of geological distribution are exactly analogous to those of geography. Closely allied species are found associated in the same beds, and the change from. species) to species appears to have been as gradual in time as in space. Geology, however, furnishes us with positive proof of the extinction and production of species, though it does not inform us how either of it has taken place. The extinction of species, however, offers but little difficulty, and the modus operandi has been well illustrated by Sir C. Lyell in his admirable ‘ Principles.’ Geological changes, however gradual, must occasionally have modified external conditions to such an extent as to have rendered the existence of certain species impossible. The extinction would in most cases be affected by a gradual dying-out, but in some instances there might have been a sudden destruction of a species of limited range. To discover how the extinct species have from time to time been replaced by new ones down to the very latest geological period, is the most difficult, and, at the same time the most interesting problem in the natural history of the earth. The present inquiry, which seeks to eliminate from known facts a law which has determined, to a certain degree, what species could and did appear at a given epoch, may, it is hoped, be considered as one step in the the right direction towards a complete solution of it.

Much discussion has of late years taken place on the question, whether the succession of life upon the globe has been from a lower to a higher degree of organization? The admitted facts seem to show that there has been a general, but not a detailed progression. Mollusca and Radiata existed before Vertebrata,