packed in the perithecia. They do not ripen in the autumn, but fall to the ground with the leaf, and there remain securely protected among the dead foliage. The following spring they mature and are liberated by the decay of the perithecia. They are then ready to attack the unfolding leaves of the willow and repeat the work of the summer before.
The wheat rust.—The development of some of the rusts, as the common wheat rust (Puccinia graminis), is even more interesting and complicated than that of the mildews. Wheat rust is also a true parasite, affecting wheat and a few other grasses. The mycelium here cannot be seen by the unaided eye, for it consists of threads which are present within the host plant, mostly in the intercellular spaces. These threads also send short branches, or haustoria (Fig. 132), into the neighboring cells to absorb nutriment.
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Fig. 280.—Sori containing Teleutospores of Wheat Rust.
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Fig. 281.—Teleutospore of Wheat Rust.
The resting-spores of wheat rust
are produced in late summer, when
they may be found in black lines
breaking through the epidermis of
the wheat stalk (black-rust stage).
They are formed in masses, called
sori (Fig. 280), from the ends of
numerous crowded mycelial strands just beneath the epidermis of
the host. The individual spores are very small and can be well
studied only with a microscope of high power
(× about 400). They are brown two-celled bodies
with a thick wall (Fig. 281). Since they are
the resting or winter-spores, they are termed teleutospores
("completed spores"). Usually they do
not fall, but remain in the sori during winter.
The following spring each cell of the teleutospore
puts forth a rather stout thread, which does not
grow more than several times the length of the
spore and terminates in a blunt extremity. This
germ tube, promycelium, now becomes divided
into four cells by cross walls, which are formed
from the top downwards. Each cell gives rise to a short, pointed
branch which, in the course of a few hours, forms at its summit
a single spore called a sporidium. This in turn germinates and
produces a mycelium. In Fig. 282 a germinating teleutospore
is drawn to show the promycelium, p, divided into four cells,
