Illustrations of Indian Botany, Vol. 1/Tamariscineae

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Illustrations of Indian Botany, Vol. 1  (1840)  by Robert Wight
Tamariscineae

XIX.-TAMARISCINEAE.

This is a small order, composed of very ramous shrubs, or small trees ; with ex-stipulate, scattered, simple, entire, sessile, minute leaves ; sometimes, in Tamarix, scale-like, closely em- bracing the stem, and lapping over each other, like tiles (imbricating.) The flowers are regu- lar, hermaphrodite, generally ranged in terminal racemes or spikes ; pedicels furnished with a small bractea.

The calyx is free or slightly adherent at the base, persistent, of 5 sepals, imbricated in (Estivation. The torus is either obsolete, or expanded into a small disk, glanduliferous on the margin. The petals equal the number of the sepals, and are alternate with them, inferior, sometimes adherent at the base, and marcescent, or withering without falling off. The stamens are hypogynous, equalling, or double, the number of the petals, rarely, by abortion, fewer than in this proportion, (this I have observed in Trichaurus ericoides,) occasionally monadelphous at the base; the anthers 2-celled, opening longitudinally. The ovary is sessile, l-celled, usually

and cannot be used as a ley for washing, as they coagulate the soap, while those growing in sweet soil in the interior are free from it. From a species or possibly a variety of T. Gallica, which grows about Mount Sinai, there exudes a kind of manna, (from the punctures of an insect which perforates its bark) which has received the name of " Manna of Mount Sinai," and has by some travellers, fancifully enough been supposed the Manna of the Scriptures. Some of the species produce abundance of galls. In Egypt the Tamarix Orientalis produces them of a deep red colour, and are much used in dyeing. All the species of this country are said by Mr. Royle to produce galls, having the properties of oak galls, but I have not been able to discover whether they are ever gathered in this part of India. It seems probable from an observation of Mr. Royle, that the galls imported into India from Mooltan, are chiefly of the Tamarix, not Oak galls, and that it is with them our bazars are principally supplied, on which account we would do well to examine the tamarix jungles, which often extend along the beds of our rivers for miles together, to ascertain to what extent they could supply our wants.

Remarks on Genera and Species. Three of the four genera referred to this order are found in India, the fourth, ( Bronnia J which seems but a doubtful member,is from America. It has been referred by Kunth to Portulacece, but is placed in this order by Dr. Lindley. The genus Trichaurus is certainly very nearly allied to Tamarix, but sufficiently distinguished by its de- candrous flowers, its cup-shaped torus bearing the stamens on its margin, and by the different form of the styles and stigmas. The beak of the seeds, which seems to form so excellent a distinguishing mark between it and Tamarix, appears to differ rather in degree than in kind, when examined under a high magnifier, since both show the beak, but in Trichaurus it is very conspicuous under the most ordinary magnifier, and even to the naked eye, while the other requires one of high powers to bring it out as represented in the figure. The dissected flower of Trichaurus is not perhaps the most suitable that might have been selected, since it seems evidently defective, in so far as having only 8 in place of 10 stamens, which last in the absence of positive evidence to the contrary, must on account of the pentasepalous calyx and 5-petaled corolla, be looked upon as the normal number. But that this is merely an irregularity, perhaps of a single flower, is rendered further probable by the circumstance of there being only 2 in place of 3 styles, three being the normal number of the order.

This figure as exhibiting a departure from the usual and regular form is not without its use, since it explains the cause of one of the greatest obstacles to the perfecting of the Linnean sexual system, depending as it does on a single set of organs, the tendency namely, of different flowers, even on the same stalk, to vary in the number of stamens and pistils, and thereby, to indicate very different places in the system for the plant to which they may belong. When our distinctive marks are taken from the relative position and number of all the different parts of the flower to each other, the formation of the ovary, and position of the ovules, added to the general habit of the plants, we have so many points of comparison, that the chances of our being misled through variations or imperfections^ in any one set of organs, are greatly diminished. Doubtless sufficient uncertainties still exist in the natural method to lead different Botanists to form different opinions both as to the situation orders ought to occupy in the series, and sometimes, in nearly allied orders as to the genera that should be respectively referred to them ; but yet, in spite of these drawbacks, its advantages over any artificial arrangement, and the Linnean is certainly the best, are such, as to ensure its general adoption by all who would study botany as a science, and not as a mere means of discovering the name of a plant, as he would the meaning of a word in a dictionary. But even this, in tropical botany, is often a very difficult operation when attempted with no other assistance than that afforded by the Sexual System, because among tropical plants, the sexual organs are so very liable to vary in number from unions among themselves, or from suppressions and additions of parts, giving rise to innumerable instances of irregular forms, among plants usually ranged in classes with regular flowers : even the Papaw, one of the most constant of dioecious plants ; I have seen with regular bi-sexual flowers.

But to return from this digression, it appears that the genus Trichaurus is amply distinguished from Tamarix by having double the number of stamens that it has sepals, and by having the filaments attached to a distinct hypogynous disk, independent of the more distinct beak of the seeds.

TAMARISCINEÆ
Illustrations of Indian Botany, Vol. 1 (page 130 crop).jpg
A. TAMARIX GALLICA. B. TRICHAURUS ERICOIDES.

The only species of the genus is, I believe, abundant on the dry banks in the bed of the Palar river below Wallajabad.

EXPLANATION OF PLATE 24.

A. 1. Tamarix gallica var Indica. Natural size. 2. Flower opened, showing the imbricated sepals, the petals, stamens, superior ovary, styles, and dilated stigmas. 3. A stamen showing the sagittate form of the anthers. 4. A single flower as it appears on being removed from the branch, the exterior pointed sepal-like leaf in front, the Bractea. 5. Ovary cut transversely, and opened, showing the parietal attachment of the ovules to the base of the carpels. 6. A mature fruit after dehiscence. 7. A seed with its downy tuft. All more or less magnified.

B. 1. Flower of Trichaurus ericoides. 2. Sepals opened, and the petals removed, to show the insertion of the stamens and ovary. 3. A stamen, anther pointed, 4. A seed with its beak. All magnified,