Page:Darwin - The various contrivances by which orchids are fertilized by insects (1877).djvu/32

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12
OPHREÆ.
Chap. I.

it is scarcely possible that any object can be pushed into it without the rostellum being touched. The exterior membrane of the rostellum then ruptures in the proper lines, and the lip or pouch is easily depressed. When this is effected, one or both of the viscid balls will almost infallibly touch the intruding body. So viscid are these balls that whatever they touch they firmly stick to. Moreover the viscid matter has the peculiar chemical quality of setting, like a cement, hard and dry in a few minutes' time. As the anther-cells are open in front, when the insect withdraws its head, or when the pencil is withdrawn, one pollinium, or both, will be withdrawn, firmly cemented to the object, projecting up like horns, as shown (fig. 2)


Fig. 2.

A. Pollen-mass of O. mascula, when first attached.
B. Pollen-mass of O. mascula, after the act of depression.


by the upper figure, A. The firmness of the attachment of the cement is very necessary, for if the pollinia were to fall sideways or backwards they could never fertilise the flower. From the position in which the two pollinia lie in their cells, they diverge a little when attached to any object. Now suppose that the insect flies to another flower, or let us insert the pencil (A, fig. 2), with the attached pollinium, into