Page:Darwinism by Alfred Wallace 1889.djvu/265

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IX
WARNING COLORATION AND MIMICRY
243

of the larvæ or some other cause, possessed disagreeable juices that caused them to be disliked by the usual enemies of their kind, they were in all probability not very different either in form or coloration from many other butterflies. They would at that time be subject to repeated attacks by insect-eaters, and, even if finally rejected, would often receive a fatal injury. Hence arose the necessity for some distinguishing mark, by which the devourers of butterflies in general might learn that these particular butterflies were uneatable; and every variation leading to such distinction, whether by form, colour, or mode of flight, was preserved and accumulated by natural selection, till the ancestral Heliconoids became well distinguished from eatable butterflies, and thenceforth comparatively free from persecution. Then they had a good time of it. They acquired lazy habits, and flew about slowly. They increased abundantly and spread all over the country, their larvæ feeding on many plants and acquiring different habits; while the butterflies themselves varied greatly, and colour being useful rather than injurious to them, gradually diverged into the many coloured and beautifully varied forms we now behold.

But, during the early stages of this process, some of the Pieridæ, inhabiting the same district, happened to be sufficiently like some of the Heliconidæ to be occasionally mistaken for them. These, of course, survived while their companions were devoured. Those among their descendants that were still more like Heliconidæ again survived, and at length the imitation would become tolerably perfect. Thereafter, as the protected group diverged into distinct species of many different colours, the imitative group would occasionally be able to follow it with similar variations,—a process that is going on now, for Mr. Bates informs us that in each fresh district he visited he found closely allied representative species or varieties of Heliconidæ, and along with them species of Leptalis (Pieridæ), which had varied in the same way so as still to be exact imitations. But this process of imitation would be subject to check by the increasing acuteness of birds and other animals which, whenever the eatable Leptalis became numerous, would surely find them out, and would then probably attack both these and their friends the Heliconidæ in order to devour