Page:Darwinism by Alfred Wallace 1889.djvu/38

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16
DARWINISM
CHAP.

wild in New Zealand. But Sir Joseph Hooker informs us that the late Mr. Bidwell habitually scattered Australian seeds during his extensive travels in New Zealand, yet only two or three Australian plants appear to have established themselves in that country, and these only in cultivated or newly moved soil.

These few illustrations sufficiently show that all the plants of a country are, as De Candolle says, at war with each other, each one struggling to occupy ground at the expense of its neighbour. But, besides this direct competition, there is one not less powerful arising from the exposure of almost all plants to destruction by animals. The buds are destroyed by birds, the leaves by caterpillars, the seeds by weevils; some insects bore into the trunk, others burrow in the twigs and leaves; slugs devour the young seedlings and the tender shoots, wire-worms gnaw the roots. Herbivorous mammals devour many species bodily, while some uproot and devour the buried tubers.

In animals, it is the eggs or the very young that suffer most from their various enemies; in plants, the tender seedlings when they first appear above the ground. To illustrate this latter point Mr. Darwin cleared and dug a piece of ground three feet long and two feet wide, and then marked all the seedlings of weeds and other plants which came up, noting what became of them. The total number was 357, and out of these no less than 295 were destroyed by slugs and insects. The direct strife of plant with plant is almost equally fatal when the stronger are allowed to smother the weaker. When turf is mown or closely browsed by animals, a number of strong and weak plants live together, because none are allowed to grow much beyond the rest; but Mr. Darwin found that when the plants which compose such turf are allowed to grow up freely, the stronger kill the weaker. In a plot of turf three feet by four, twenty distinct species of plants were found to be growing, and no less than nine of these perished altogether when the other species were allowed to grow up to their full size.[1]

But besides having to protect themselves against competing plants and against destructive animals, there is a yet deadlier