Page:Darwinism by Alfred Wallace 1889.djvu/396

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372
DARWINISM
CHAP.

or perish, and all which come within sight of an island will struggle to reach it as their only refuge. But, with mountain summits the case is altogether different, because, being surrounded by land instead of by sea, no bird would need to fly, or to be carried by the wind, for several hundred miles at a stretch to another mountain summit, but would find a refuge in the surrounding uplands, ridges, valleys, or plains. As a rule the birds that frequent lofty mountain tops are peculiar species, allied to those of the surrounding district; and there is no indication whatever of the passage of birds from one remote mountain to another in any way comparable with the flights of birds which are known to reach the Azores annually, or even with the few regular migrants from Australia to New Zealand. It is almost impossible to conceive that the seeds of the Himalayan primula should have been thus carried to Java; but, by means of gales of wind, and intermediate stations from fifty to a few hundred miles apart, where the seeds might vegetate for a year or two and produce fresh seed to be again carried on in the same manner, the transmission might, after many failures, be at last effected.

A very important consideration is the vastly larger scale on which wind-carriage of seeds must act, as compared with bird-carriage. It can only be a few birds which carry seeds attached to their feathers or feet. A very small proportion of these would carry the seeds of Alpine plants; while an almost infinitesimal fraction of these latter would convey the few seeds attached to them safely to an oceanic island or remote mountain. But winds, in the form of whirlwinds or tornadoes, gales or hurricanes, are perpetually at work over large areas of land and sea. Insects and light particles of matter are often carried up to the tops of high mountains; and, from the very nature and origin of winds, they usually consist of ascending or descending currents, the former capable of suspending such small and light objects as are many seeds long enough for them to be carried enormous distances. For each single seed carried away by external attachment to the feet or feathers of a bird, countless millions are probably carried away by violent winds; and the chance of conveyance to a great distance and in a definite direction must be many