all the year round, at 14,000 ft.; Sincholagua had large beds of permanent snow at 15,300 ft., Cotopaxi was permanently covered at 15,500 ft. on its western side, Corazon had daily snowstorms down to 14,500 ft., but no permanent beds of snow on its east side (elevation 15,871 ft.); and Chimborazo had deep snow at 15,600 ft. on its north-east and south sides in June—July. The eastern range was found to receive the heaviest snowfall. The elevation at which human residence is possible seems to be unusually high in Ecuador. Many of the towns and villages of central Ecuador lie at altitudes ranging from 8606 ft. (Ambato) to 9839 ft. (Machachi). The capital city of Quito is 9343 ft. above the sea, and is celebrated for its agreeable temperature, and also for its healthiness in spite of prevailing unsanitary conditions. Above these towns are a number of farms and herdsmen’s habitations, where men live the whole or a part of the year with less discomfort from low temperature than is experienced in northern Europe and northern United States. According to Whymper, the tambo of Chuquipoquio, at the foot of Chimborazo, is 11,704 ft., and the hacienda of Pedregal, near Iliniza, 11,629 ft., both being permanently occupied. The hacienda of Antisana, 13,306 ft., and the herdsmen’s hut of Cunayaco on Chimborazo, 13,396 ft., are occupied only for a part of the year. The highest elevations are generally covered with ice and snow, and glaciers, according to Whymper, are to be found upon no less than nine of the culminating peaks, and possibly upon two or three more. These serve to modify the temperatures of the plateau, which is swept by cold winds at all seasons of the year. The prevailing wind is that of the north-east and south-east trade winds, broken and modified on the plateau and western lowlands by mountain barriers. Westerly and north-west winds are sometimes experienced, but are not permanent.
Flora.—The flora of the Quito basin has been well studied by various European botanists, more especially by Dr William Jameson (1796–1873) of the university of Quito, who began the preparation of a synopsis of the Ecuadorean flora in 1864–1865 (Synopsis plantarum Quitensium, 2 vols., Quito, 1865). The flora of the forested lowlands on both sides of the Andes has not been studied and described so fully. From the Pacific coast upward to a height of about 3000 to 4000 ft. the vegetation is distinctively tropical, including among its economic products cacao, cotton, sugar, tobacco, rice, maize, yucca (also known as cassava and mandioca), peanuts, bananas, sweet potatoes, yams, arracacha (Conium moschatum, H.B.K., or Arracacha esculenta), indigo, rubber (Castilloa), ivory-nuts, cinchona and bread-fruit. Most of these become rare at 3000 ft., but a few, like sugar-cane, are cultivated as high as 8000 ft. The alluvial valley of the Guayas, above Guayaquil, is celebrated for the richness of its vegetation, which, in fruit alone, includes cacao, coffee, coco-nuts, pine-apples, oranges, lemons, guayavas (Psidium pomiferum), guavas (Inga spectabilis), shaddocks (or grape-fruit), pomegranates, apricots, chirimoyas (Anona Chirimolia), granadillas (Passiflora quadrangularis), paltas (Persea gratissima, otherwise known as “alligator pears”), tunas (Cactus), mangoes (Mangifera Indica), pacays (Prosopis dulcis), aji (Chile pepper), and many others of less importance. Besides rubber, the forests produce a great variety of cabinet and construction woods, ivory-nuts (from the “tagua” palm, Phytelephas macrocarpa), “toquilla” fibre (Carludovica palmata) for the manufacture of so-called Panama hats, cabbage palms, several species of cinchona, vanilla and dyewoods. Among the large trees which are valued for their timber are redwood (Humiria balsamifera), Brazil-wood, algarrobo, palo de cruz (Jacquinea ruscifolia), guaiacum or holy wood, rosewood, cedar and walnut. From 6000 to 10,000 ft. above the sea, the indigenous species include the potato, maize, oca (Oxalis tuberosa), and quinua (Chenopodium quinoa), and the exotic species, wheat, barley, oats, alfalfa (Medicago sativa), and most of the fruits and vegetables of the northern temperate zone. Wheat does not form a head below 4500 ft., nor ripen above 10,500. The larger forest trees are rarely seen above 10,000 ft., and even there only on the outer slopes of the Cordilleras. The Escallonia myrtalloides, however, is found at an elevation of 13,000 ft., and the shrubby Befarias 400 or 500 ft. higher. A characteristic growth of the open plateau and upland valleys is the cabulla, cabaya or maguey (Agave americana), whose fibre is much used by the natives in the manufacture of cordage, sandals (alpargatas) and other useful articles. In the treeless region lying between 11,600 and 13,800, or in other places between 12,000 and 14,000 ft., the similarity of the vegetation to that of the corresponding European region, according to Wagner, is especially striking. On the paramos of Chimborazo, Pichincha, Iliniza, &c., the relation of characteristic genera to those identical with genera in the Alpine flora of Europe is as 5 to 4; and the botanist might almost suppose himself in the Upper Engadine. Of the flora of the highest Andes, Whymper found 42 species, of various orders, above 16,000 ft., almost all of which were from Antisana and Chimborazo; 12 genera of mosses were found above 15,000 ft., and 59 species of flowering plants above 14,000 ft., of which 35 species came from above 15,000 and 20 species from above 16,000 ft. The highest specimen obtained was a lichen (Lecanora subfusca, L.) on the south side of Chimborazo, 18,400 ft. above sea-level. Mosses (Grimmia) were found on Chimborazo at 16,660 ft., ferns (Polypodium pycnolepis, Kze.) at 14,900, and specimens of Gentiana rupicola, H. B. K., Achyrophorus quitensis, Sz. Bip., Culcitium nivale, H. B. K., at 16,300; Phyllactis inconspicua, Wedd., at 16,600, Astragalus geminiflorus, H. B. K., at 14–15,000, Geranium diffusum, H. B. K., at 16,000, Malvastrum phyllanthos, Asa Gray, at 16,500, Draba obovata, Benth., at 16,660, and Ranunculus praemorsus, Kth., at 16,500—all on Chimborazo. Fuchsia loxensis, H. B. K., was found on the slope of Sara-Urcu at 12,779 ft., and currant bushes (Ribes glandulosum, R. & P.), on Chimborazo, at 14,000. On the eastern slopes of the Andes, where the rainfall is continuous throughout the year and the atmosphere is surcharged with moisture, the forest growth is phenomenal. It is similar to that of the Colombian and Peruvian montanas, modified, if at all, by the excessive humidity which prevails in this region.
Fauna.—The fauna of Ecuador is comparatively poor in mammalia, but the birds and still more the insects are very numerous. The Quadrumana are represented by a large number of species, the eastern forests being very much like the other parts of the great Amazonian basin in this respect. The Carnivora include the puma (Felis concolor), jaguar (F. onca), ocelot (F. grisea), bear (Ursus ornatus), fox, weasel and otter. A small deer and, in southern Ecuador, the llama (Auchenia) with its allied species, the alpaca, guanaco and vicuña, represent the ruminants. The rodents are numerous and include most, if not all, of the Amazonian species—the capybara (Hydrochoerus capybara), cavia (C. aperea), paca (Coelogenys paca) and cutia (Dasyprocta aguti), all amphibious and having an extensive range. Tapirs are to be found in the eastern forests, the peccary in more open woodlands, and the opossum in nearly every part of the country. Cattle, horses, asses, sheep and swine were introduced by the Spaniards, and thrive well in some of the provinces. Excellent horses are reared in the uplands, as well as mules and cattle, the pasturage on the mountain slopes being good, and alfalfa being grown in abundance in many districts. The Reptilia include countless numbers of alligators in the Guayas and its tributaries and in the tide-water channels of many of the smaller rivers; many species of lizards, of which Mr Whymper found three in the Quito basin; snakes of every description from the huge anaconda of the Amazon region down to the beautifully marked coral snake; and a great variety of frogs and toads. Bats also are very numerous, especially in the eastern forest region, where the vampire bat is a serious obstacle to permanent settlement. The avifauna of Ecuador is distinguished for the great variety of its genera and species, among which are many peculiar to the Amazon valley, and others to the colder uplands. Among the Amazon species may be mentioned the parrot, macaw (Macrocercus), toucan (Ramphastos), curassow (Crax), penelope, trogon, and horned screamer (Palamedea cornuta). There are also herons, ibises, storks and cranes, including the great black-headed white crane, Mycteria americana, which ranges from northern Argentina to Colombia. One species of ibis, the Theristicus caudatus, is to be found, it is said, only on the slopes of Antisana. Species of the pheasant and partridge are not uncommon, and the “guácharo” (Steatornis caripensis), once believed to inhabit Venezuela only, is found in Ecuador also. The Raptores are well represented by a large number of genera and species, which include the condor, eagle, vulture, falcon, hawk and owl. The condor (Sarcorhamphus gryphus) is commonly found between the elevations of 6000 and 16,000 ft., rarely, if ever, descending to the lowland plains or rising above the lower peaks. It preys upon the smaller animals and inflicts much loss upon stock farmers through the destruction of calves, lambs, &c., but it very rarely ventures to attack man or any of the larger animals. The eagle common to Ecuador is the Morphnus taeniatus, and possibly the M. guaianensis on the eastern slopes of the Andes. The harrier-eagle (Herpetotheres cachinnans) is also to be found throughout this part of the continent. An eagle with buzzard-like habits, the Leucopternis plumbea, is likewise common in Ecuador. Among the vultures the turkey-buzzard group (Rhinogryphus or Cathartes), including the R. aurus, burrovianus and perniger, is common everywhere. The carrion crow, or black vulture (Catharista atrata), is also common to every part of the country, and is the general scavenger. The carrion hawks are represented by the Polyborus tharus, popularly called the “caracara,” and the Phalcobaenus carunculatus; the falcons by the Aesalon columbarius; and the kites by the Gampsonyx swainsoni. The Ecuadorean owl is the Bubo nigrescens. An interesting species of the song birds is popularly known as the “flautero” (flute-bird), which inhabits the eastern forests. Its notes are marvellous imitations of “the most mellow, sweet-sounding flute,” but the singer itself, according to Mr Simson, is “a very insignificant-looking little, greyish-coloured bird,” which “always dies in captivity.” The most interesting group of the smaller birds is that of the hummingbirds, of which the number and variety is astonishing. Some of these have a very wide range, while others are apparently limited to a small district, or to a certain altitude. The best-known fish of Ecuador is the insignificant Pimelodus cyclopum, the only fish found in the streams and lakes of the plateau region. Its fame rests on Humboldt’s publication of the tradition that great numbers of this tiny fish had been thrown out during the eruptions of Imbabura and other volcanoes. Mr Whymper’s explanation of the phenomenon is that the fish are scattered over the land by the sudden overflow during volcanic eruptions of the rivers and lakes which they inhabit. The rivers of the eastern plains are probably stocked with the fish found in the Amazon. On the coast, the Ancon de Sardinas bay is so named from the multitude of small fish (sardinas) which inhabit its