Page:History of botany (Sachs; Garnsey).djvu/263

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Chap. I.]
by Malpighi and Grew.
243

himself states incidentally, that he lit upon this notion from looking at shrivelled masses of tissue, when he naturally saw wrinkles and folds, which he took for threads. Besides he seems to have used blunt knives, which might easily tear the cell-walls into threads; so we might gather from the figure in Plate 40, where what he supposes to have been a tissue of thread from the walls of a cell is depicted quite plainly. Lastly the observation of vessels with reticulated thickening, and parenchyma-cells with crossed striation may have contributed to his view.

It will hardly be superfluous to remark here, that Grew's idea of this very delicate structure of cell-walls has evidently given rise to the common expression cell-tissue (contextus cellulosus) when speaking of plants and animals, an expression which has become naturalised in microscopy, and is still retained though we no longer think of Grew's comparison of cell-structure with artificial lace. But the word tissue has often misled later writers, as words are apt to do, and made them found their conception of vegetable structure on the resemblance to an artificial tissue of membranes and threads.

Grew, like Malpighi, derives the young layers of wood in the stem from the innermost layers of the rind. The true wood, he says on page 114, is entirely composed of old lymph-vessels, that is of fibres, which lay originally in the inner circumference of the rind. But by true woody substance he understands the fibrous components of the wood, excluding the air-vessels; his lymph-vessels are the bast-fibres and similar forms; for, he goes on, the air-vessels with the medullary rays and the true wood form what is commonly called the wood of a tree; he uses the term air-vessels, not because these forms never contain sap, but because they only contain a vegetable air during the proper period of vegetation, when the vessels of the rind are filled with sap.

The above is certainly a very imperfect account of Grew's services to phytotomy; for the points here made prominent