Page:History of botany (Sachs; Garnsey).djvu/69

From Wikisource
Jump to navigation Jump to search
This page has been validated.
Chap. II.]
from Cesalpino to Linnaeus.
49

quence of the Linnaean terminology, which neglected all comparative examination.

Moreover the doctrine of metamorphosis appears in a more consistent and necessary form in Cesalpino than in the botanists of the 19th century before Darwin; it flows more immediately from his philosophical views on the nature of plants, and appears therefore up to a certain point thoroughly intelligible. We may also consider as part of this doctrine in Cesalpino the view that the substance of the seed (embryo and endosperm) arises from the pith, because the pith contains the vital principle[1], and as the pith in the shoot is surrounded for protection by the wood and the bark, so the substance of the seed is surrounded by the woody shell, and by the bark-like pericarp or by a fruit-envelope answering to a pericarp. According to Cesalpino therefore the substance of the seed with its capability of development springs from the pith, the woody shell from the wood, the pericarp from the rind of the shoot. The difficulty which arises from this interpretation, namely, that in accordance with his theory the parts of the flower also, the calyx, the corolla, and the stamens ought to spring from the outer tissues of the shoot, he puts aside with the remark (p. 19) that these parts of the flower are formed when the pericarp is still in a rudimentary state; that the pericarp is only fully developed after these parts have fallen off, and that they are so thin that there is nothing surprising in this view of the matter. We see in Cesalpino's doctrine of metamorphosis without doubt the theory of the flower afterwards adopted by Linnaeus, though in a somewhat different form. That Linnaeus himself regarded the theory ascribed to him on the nature of the flower


  1. We find it stated in Theophrastus that if the pith of the vine is destroyed the grapes contain no stones; this evidently points to a still higher antiquity for the view that the seeds are formed from the pith; see the De causis plantarum, v. ch. 5, in the ' Theophrasti quae supersunt opera' of Schneider, Leipzig, 1818.