Page:Insectivorous Plants, Darwin, 1899.djvu/82

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when leaves were immersed in water, at a lower temperature than 130° (54°.4 Cent.), which was then raised to 145° (62°.7 Cent.), they became during the earlier period of immersion inflected, but on being afterwards left in cold water were incapable of re-expansion. Exposure for a few minutes to a temperature of 145° sometimes causes some few of the more sensitive glands to be speckled with the porcelain-like appearance; and on one occasion this occurred at a temperature of 140° (60° Cent.). On another occasion, when a leaf was placed in water at this temperature of only 140°, and left therein till the water cooled, every gland became like porcelain. Exposure for a few minutes to a temperature of 150° (65°.5 Cent.) generally produces this effect, yet many glands retain a pinkish colour, and many present a speckled appearance. This high temperature never causes true inflection; on the contrary, the tentacles commonly become reflexed, though to a less degree than when immersed in boiling water; and this apparently is due to their passive power of elasticity. After exposure to a temperature of 150° Fahr., the protoplasm, if subsequently subjected to carbonate of ammonia, instead of undergoing aggregation, is converted into disintegrated or pulpy discoloured matter. In short, the leaves are generally killed by this degree of heat; but owing to differences of age or constitution, they vary somewhat in this respect. In one anomalous case, four out of the many glands on a leaf, which had been immersed in water raised to 156° (68°.8 Cent.), escaped being rendered porcellanous ;[1] and the proto-plasm in the cells close beneath these glands underwent some slight, though imperfect, degree of aggregation.

Finally, it is a remarkable fact that the leaves of Drosera rotundifolia, which flourishes on bleak upland moors throughout Great Britain, and exists (Hooker) within the Arctic Circle, should be able to withstand for even a short

time immersion in water heated to a temperature of 145°.[2]

  1. As the opacity and porcelain-like appearance of the glands is probably due to the coagulation of the albumen, I may add, on the authority of Dr. Burton Sanderson, that albumen coagulates at about 155°, butn in presence of acids, the temperature of coagulation is lower. The leaves of Drosera contain an acid, and perhaps a difference in the amount contained may account for the slight differences in the results above recorded.
  2. It appears the cold-blooded animals are, as might have been