Abstract
The species includes 39 subspecies. The type locality is Georgia. Seven populations of D. (C.) s. scabricolle are known from the republic. One population of the type subspecies is situated in north Armenia.
D. (C.) s. shirakense, ssp. n. (type locality Lernavan environs, 40°48′10.57″N 44°09′58.10″E, 1893 m) is distributed all over west part of Armenia and in north-east Turkey.
Seven more subspecies are recognized in Armenia: D. (C.) s. tavushense, ssp. n. (type locality–Idzhevan environs, Acharkut, about 41°2′9″N, 45°5′6″E, 800 m) in north Armenia, D. (C.) s. gegarkunicum, ssp. n. (type locality–Akhberk environs, 40°32′58″N, 45°17′09″E, 2280 m) around north part of Sevan Lake, D. (C.) s. sevangense Reitter, 1889 (type locality–environs of Sevan-city)–high mountains all acrros cenral part of Armenia, D. (C.) s. pseudosevangense, ssp. n. (type locality–Noravan environs, 39°32′15″N, 46°07′56″E, 2000 m) at the east part of D. (C.) s. sevangense area, D. (C.) s. vaykense, ssp. n. (type locality–Vayk environs, 3.6 km northwards Zaritap, 39°40′11″N, 45°30′28″E, 1300 m)–south of Marz Vayots Dzor, D. (C.) s. tekhense, ssp. n. (type locality–Tekh, 39°34′6″N, 46°25′52″E, 1600 m)–north-east of Marz Syunik, D. (C.) s. tuzovi, ssp. n. (type locality–Megri environs)–south-east part of Armenia with penetration to Nakhchivan Autonomous Republic; a new subspecies is described from Nagorno-Karabakh Republic: D. (C.) s. artsakhense, ssp. n. (type locality–Shishkend environs, 39°45′52″N, 46°48′09″E, 1190 m).
Six subspecies are accepted for Nakhchivan Autonomous Republic including D. (C.) s. buzgoviense, ssp. n. (type locality–Buzgov environs, about 39°31′7″N, 45°23′23″E, 1700 m), D. (C.) s. shakhbuzum, ssp. n. (type locality–Shakhbuz District, Bichenek environs, 39°30′50″N, 45°46′28″E, 1940 m), D. (C.) s. araxense, ssp. n. (type locality–Negram environs, 39°5′N, 45°24′13″E, 790 m), D. (C.) s. paiz Danilevsky, 1999 (type locality–Paiz env., 39°25′30″N, 45°23′14″E, 1300 m), D. (C.) s. nakhiczevanum Danilevsky, 1999 (type locality-south slope of Bichenek pass, 39°31′58″N, 45°47′51″E, 2100 m), D. (C.) s. dobrovljanskii Suvorov, 1915 (type locality–Kyukyu village, about 39°31′27″N, 45°37′18″E).
Two subspecies are distributed in Azerbaijan: D. (C.) s. modestum Tournier, 1872, nom. rest. (type locality–Azerbaijan, Shamkir [before Annenfeld], 40°50′83″N, 46°01′02″E)–from north-west Azerbaijan and
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