Page:Makers of British botany.djvu/280

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MILES JOSEPH BERKELEY

and consist mainly of critical notes on the morphology and affinities of the fungi under consideration, and will compel the attention of mycologists for all time.

From the above brief account it may perhaps be concluded that Berkeley was essentially a systematist and founder of new species. Owing to the vast amount of material that passed through his hands, he was so perforce, but his leaning was always rather towards the biological and morphological side of the subject.


Morphology and Literature.

The first important paper dealing with the morphology of the hymenial structure in Fungi, is entitled, "On the Fructification of the Pileate and Clavate Tribes of the Hymenomycetous Fungi," Annals of Nat. Hist., 1838. Here is clearly demonstrated for the first time, the universal occurrence of basidia bearing spores at their summit, throughout the entire group of fungi known to-day as the Hymenomycetes, including Agaricaceae, Thelephoraceae, Clavariaceae, etc. This important discovery rendered possible the basis of a classification on morphological grounds, which holds good at the present day. A careful study of the text and illustrations demonstrates the fact that Berkeley was perfectly well acquainted with all the essential details of the hymenium, many of which have been repeatedly rediscovered and described under new names, in ignorance of the fact that such structures had previously been equally well described.

Berkeley continued his investigations on the structure of the hymenium, and his next paper, entitled "Sur la fructification des genres Lycoperdon, Phallus et de quelques autres genres voisins," in Annal. Sci. Nat. Ser. 2, vol. XII. (1839), demonstrated the universal presence of basidia bearing spores at their summit in the family now known as the Gasteromycetes. This research on the part of Berkeley led to the universal adoption of the two primary divisions of the Fungi; Basidiomycetes, having the spores borne at the apex of a basidium; and Ascomycetes, having the spores produced within specialised sacs, or asci.