Page:Miscellaneousbot01brow.djvu/309

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OF CENTRAL AFRICA.
291

fore, exceptions to all the assumed characters of Mimoseæ are found, and there is some approach in both genera to the habit of Cæsalpineæ. It is still possible, however, to distinguish, and it will certainly be expedient to preserve, those two tribes or orders. Abandoning divisions strictly natural, and so extensive as the tribes in question, merely because we may not be able to define them with precision, while it would imply, what is far from being the case, that our analysis of their structure is complete, would, at the same time, be fatal to many natural families of plants at present admitted, and among others to the universally received class to which these tribes belong. No clear character, at least, is pointed out in the late elaborate work of M. De Candolle,[1] by which Leguminosæ may be distinguished from Terebintaceæ and Rosaceæ, the orders supposed to be most nearly related to it. It is possible, however, that such characters, though hitherto overlooked, may really exist; and I shall endeavour to show that Leguminosæ, independent of the important but minute differences in the original structure and development of its ovulum, may still be distinguished at least from Rosaceæ.

In the character of Polygaleæ, which I published in 1814,[2] I marked the relation of the parts of the floral envelopes to the axis of the spike, or to the subtending bractea. I introduced this circumstance chiefly to contrast Polygaleæ with Leguminosæ, and to prove, as I conceived, that Securidaca, which had generally been referred to the latter family, really belonged to the former.

M. De Jussieu, who soon after published a character of Polygaleæ, entirely omitted this consideration, and continued to refer Securidaca to Leguminosæ. M. De Candolle, however, in the first volume of his 'Prodromus,' has adopted both the character and limits of Polygaleæ, which I had proposed, though apparently not altogether satisfied with the description he himself has given of the divisions of the calyx and corolla.

The disposition of the parts of the floral envelopes, with

  1. Mémoires sur la Famille des Legumineuses.
  2. Flinders's Voy. to Terra Anstr. 2, p. 542. (Antè, pp. 13, 14.)