Page:Onthetendencyofspecies.pdf/4

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48
MESSRS. C. DARWIN AND A. WALLACE ON THE

necessarily stop as soon as the country was fully stocked; and yet we have every reason to believe, from what is known of wild animals, that all would pair in the spring. In the majority of cases it is most difficult to imagine where the checks fall—though generally, no doubt, on the seeds, eggs, and young; but when we remember how impossible, even in mankind (so much better known than any other animal), it is to infer from repeated casual observations what the average duration of life is, or to discover the different percentage of deaths to births in different countries, we ought to feel no surprise at our being unable to discover where the check falls in any animal or plant. It should always be remembered, that in most cases the checks are recurrent yearly in a small, regular degree, and in an extreme degree during unusually cold, hot, dry, or wet years, according to the constitution of the being in question. Lighten any check in the least degree, and the geometrical powers of increase in every organism will almost instantly increase the average number of the favoured species. Nature may be compared to a surface on which rest ten thousand sharp wedges touching each other and driven inwards by incessant blows. Fully to realize these views much reflection is requisite. Malthus on man should be studied; and all such cases as those of the mice in La Plata, of the cattle and horses when first turned out in South America, of the birds by our calculation, &c., should be well considered. Reflect on the enormous multiplying power inherent and annually in action in all animals; reflect on the countless seeds scattered by a hundred ingenious contrivances, year after year, over the whole face of the land; and yet we have every reason to suppose that the average percentage of each of the inhabitants of a country usually remains constant. Finally, let it be borne in mind that this average number of individuals (the external conditions remaining the same) in each country is kept up by recurrent struggles against other species or against external nature (as on the borders of the Arctic regions, where the cold checks life), and that ordinarily each individual of every species holds its place, either by its own struggle and capacity of acquiring nourishment in some period of its life, from the egg upwards; or by the struggle of its parents (in short-lived organisms, when the main check occurs at longer intervals) with other individuals of the same or different species.

But let the external conditions of a country alter. If in a small degree, the relative proportions of the inhabitants will in most cases simply be slightly changed; but let the number of