Page:Popular Science Monthly Volume 3.djvu/471

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THE PROBLEMS OF THE DEEP SEA.
457

region they distinguish a fifth, which is never uncovered, and is inhabited by oysters, scallops, and large starfishes and other animals. Beyond this they seem to think that animal life is absent.[1]

Audouin and Milne Edwards were the first to see the importance of the bearing of a knowledge of the manner in which marine animals are distributed in depth, on geology. They suggest that, by this means, it will be possible to judge whether a fossiliferous stratum was formed upon the shore of an ancient sea, and even to determine whether it was deposited in shallower or deeper water on that shore; the association of shells or animals which live in different zones of depth will prove that the shells have been transported into the position in which they are found; while, on the other hand, the absence of shells in a deposit will not justify the conclusion that the waters in which it was formed were devoid of animal inhabitants, inasmuch as they might have been only too deep for habitation.

The new line of investigation thus opened by the French naturalists was followed up by the Norwegian, Sars, in 1835, by Edward Forbes, in our own country, in 1840,[2] and by Œrsted, in Denmark, a few years later. The genius of Forbes, combined with his extensive knowledge of botany, invertebrate zoology, and geology, enabled him to do more than any of his compeers in bringing the importance of distribution in depth into notice; and his researches in the Ægean Sea, and still more his remarkable paper "On the Geological Relations of the Existing Fauna and Flora of the British Isles," published in 1846, in the first volume of the "Memoirs of the Geological Survey of Great Britain," attracted universal attention.

On the coasts of the British Islands, Forbes distinguishes four zones or regions, the Littoral (between tide-marks), the Laminarian (between low-water mark and 15 fathoms), the Coralline (from 15 to 50 fathoms), and the Deep sea or Coral region (from 50 fathoms to beyond 100 fathoms). But, in the deeper waters of the Ægean Sea, between the shore and a depth of 300 fathoms, Forbes was able to make out

  1. "Enfin plus bas encore, c'est-à-dire alors loin des côtes, le fond des eaux ne paraît plus être habité, du moins dans nos mers, par aucun de ces animaux" (l. c, tome i., p. 237). The "ces animaux" leaves the meaning of the authors doubtful.
  2. In the paper in the "Memoirs of the Survey" cited farther on, Forbes writes:

    "In an essay 'On the Association of Mollusca on the British Coasts, considered with reference to Pleistocene Geology,' printed in the Edinburgh Academic Annual for 1840, I described the mollusca, as distributed on our shores and seas, in four great zones or regions, usually denominated 'The Littoral Zone,' 'The region of Laminariæ,' 'The region of Corallines,' and 'The region of Corals.' An extensive series of researches, chiefly conducted by the members of the committee appointed by the British Association to investigate the marine geology of Britain by means of the dredge, have not invalidated this classification, and the researches of Prof. Lovén, in the Norwegian and Lapland Seas, have borne out their correctness. The first two of the regions above mentioned had been previously noticed by Lamouroux, in his account of the distribution (vertically) of sea-weeds, by Audouin and Milne Edwards in their 'Observations on the Natural History of the Coast of France,' and by Sars in the prefaoe to his 'Beskrivelser og Jagttagelser.'"