which give the results of a portion of the actual experiments. As the experiments were rather extensive, it is not possible here to present anything like the complete material. Only representative matings are here given. Table II. gives the results of some of the B. P. R. × B. P. B. matings in detail, in order to show, not only the accordance between observation and theory, but also the distinctness of the classes of fecundity segregated (shown by the average winter production in each segregated class).
From the data set forth in the above table there can be no doubt as to the fact of the Mendelian segregation of fecundity, nor as to the entire distinctness of the things segregated.
In order to give a general survey of the results, and to demonstrate the reality of segregation over the wide range of material included in the experiments, the summary Table III is presented.
Showing the Observed and Expected Distribution in Respect of Fecundity of the Adult Female Offspring from all Matings in each of the Classes Tested in the Experiments
|Matings||Winter Production of Daughters|
|Class||Over 30||Under 30||Zero|
|All Barred Plymouth Rock × Barred||Observed||365||.5||259||.5||31|
|All Cornish Indian Game × Indian||Observed||2||23||15|
|All F1 (B.P.R. × C.I.G. and reciprocal||Observed||36||79||8|
|All F2 (F1 × F1, and F1 × parent forms||Observed||57||.5||98||.5||23|
|in all possible combinations).||Expected||68||.60||95||.00||15||.40|
Considering the nature of the material and the character dealt with the agreement shown between observation and hypothesis is certainly as close as could reasonably be expected. Such discrepancies as are shown in the above table are fully discussed and their probable physiological explanations set forth in detail in the complete account of these experiments.
The detailed data given in the complete paper, of which the above discussion and tables give merely a very incomplete abstract, appear definitely to establish the following points:
1. That fecundity in the domestic fowl is inherited strictly in accordance with Mendelian principles.
2. That observed individual variations in fecundity here depend upon two separately inherited physiological factors, L1 and L2.
3. That high fecundity is manifested only when both of these factors are present together in the same individual.
4. That either of these factors when present alone whether in homo-