Page:Principal Characters of American Jurassic Dinosaurs; Part VI, Restoration of Brontosaurus.pdf/3

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O. C. Marsh—Restoration of Brontosaurus.
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place, they are attached to the occiput just above the foramen magnum, and extend backward and outward, overlapping the lateral pieces of the atlas, thus protecting the spinal cord at this point, which would otherwise be much exposed.

These bones are short, flattened, and slightly curved, resembling somewhat a riblet. The anterior end is thickened and rugose for attachment to a roughened surface on the exoccipital, just above and outside the foramen magnum. The shaft is flattened from above downward, and gradually converges to a thin posterior end. In Morosaurus grandis, these bones are about 65 mm. in length, and 30 along the surface which joins the occiput. They correspond in position to the muscle in mammals known as the rectus capitis posticus minor.

In the existing Cormorants (Graculus) a single slender bone is articulated to the occiput on the median line. This, however, does not correspond to the bones here described. To distinguish it from the post-occipitals, it may be called the nuchal bone.

Stapes wanting.

In the skull of Morosaurus in which the post-occipital bones were found in position and the other bones at the base of the skull were undisturbed, a careful search was made for the stapes, but no indication of it was found. Its absence in this specimen, so well preserved, would indicate that it was wanting in this genus, if not in the other Sauropoda.

Columella present.

In a skull of Brontosaurus in which the bones, although displaced, were in very perfect preservation, a pair of bones were found which apparently are the columellæ. They are elongated, flattened bones, with the shaft somewhat constricted in the middle, and twisted. Their length corresponds to the elevated posterior part of the skull in this genus.

Hyroid bones.

There are two pairs of hyoid bones in the Sauropoda. They are elongated, rodlike, and somewhat curved. In Brontosaurus excelsus, they are 210 and 130 mm. in length respectively.

Among the other points of interest in the skull of the Sauropoda are the following:

The parietal bones are very short, and form but a small portion of the brain case. They are composed chiefly of the flattened arched processes, which meet with the squamosals at their outer ends. There is no parietal foramen. The squamosals lie upon the par-occipital processes. They have a