Page:Scientific results HMS Challenger vol 18 part 1.djvu/262

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54
THE VOYAGE OF THE H.M.S. CHALLENGER.

is a medullary shell. In this case the size of the capsule remains intermediate between the inner and the middle shell.

The Tetrasphærida, or the Sphæroidea with four concentric shells, are in general not frequent, and not rich in different forms. In most of the observed species two inner shells are medullary, two outer cortical shells, the former within, the latter without, the central capsule; and the wall of the capsule, pierced by the connecting radial beams, lies between the two middle shells. But there are some Tetrasphærida in which all four shells seem to be external or cortical shells.

The Polysphærida, or the Sphæroidea with five or more concentric shells, seem of course to offer the greatest possibility for the development of very different forms; but in reality this group is the poorest and smallest of all; and only one part of it, the Arachnosphærida, is rather common. In this peculiar division the shell is composed of five to ten or more, very delicate, cobweb-like concentric shells, which are connected by radial beams; all are cortical shells, and lie outside the central capsule. Much more rare are those Polysphærida, in which both innermost shells, as true medullary shells, lie within the central capsule, all others being outside it. The total number of concentric shells in this group is commonly between five and ten, rarely more.

The Spongosphærida are distinguished from all other Sphæroidea by the spongy structure of the spherical shell, which is composed wholly or partially of an irregular spongy framework. The relation of this group to the other groups of Sphæroidea is probably rather complicated, for in some Spongosphærida the whole shell is composed of massive spongy reticulation, whilst in others it contains a spherical central cavity, and in a third group this cavity is filled up by one or two concentric lattice-shells, connected by radial beams. Many of these Spongosphærida are very common, and of considerable size.

The Collosphærida form a peculiar separate group of Sphæroidea, distinguished from all others by their social life or aggregation in colonies (cœnobia). They represent the only group of Sphærellaria in which this association of numerous individual capsules or cells is realised. The shell is almost constantly simple, without regularly disposed radial spines; therefore they may be called "social Monosphærida," or better "polyzoic Ethmosphærida." Only in one small group (Clathrosphærida) the shell, enveloping every central capsule, is double or surrounded by an external mantle; these may be compared to the Diplosphærida (or better to a part of the Carposphærida, Liosphæra, p. 76). In most of the Collosphærida the lattice-shell is more or less irregular in form and structure.


The Lattice Work of the fenestrated shells is in the Sphæroidea of the greatest variability, and its innumerable modifications serve mainly for the distinction of species. In general we can distinguish as the most important modifications a regular network (with equal size, form, and distance of the pores or meshes) and an irregular network (with