Page:The Formation of Vegetable Mould through the Action of Worms (1881).djvu/260

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246
DISINTEGRATION
Chap. V.

therefore necessary to obtain flints for building purposes from the bed of red clay overlying the chalk (the residue of its dissolution by rainwater) or from the chalk itself.

Not only do worms aid indirectly in the chemical disintegration of rocks, but there is good reason to believe that they likewise act in a direct and mechanical manner on the smaller particles. All the species which swallow earth are furnished with gizzards; and these are lined with so thick a chitinous membrane, that Perrier speaks of it,[1] as "une véritable armature." The gizzard is surrounded by powerful transverse muscles, which, according to Claparède, are about ten times as thick as the longitudinal ones; and Perrier saw them contracting energetically. Worms belonging to one genus, Digaster, have two distinct but quite similar gizzards; and in another genus, Moniligaster, the second gizzard consists of four pouches, one succeeding the other, so that it may almost be said to have five gizzards.[2] In the same

  1. 'Archives de Zoolog. expér.' tom. iii. 1874, p. 409.
  2. 'Nouvelles Archives du Muséum,' tom. viii. 1872, p. 95, 131.