Page:The Formation of Vegetable Mould through the Action of Worms (1881).djvu/67

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Chap. I.
CALCIFEROUS GLANDS.
53

with fine ferruginous sand, it was manifest that the oxide of iron, with which the grains of silex were coated, had been dissolved and removed from them in the castings.

The digestive fluid of worms resembles in its action, as already stated, the pancreatic secretion of the higher animals; and in these latter, "pancreatic digestion is essentially alkaline; the action will not take place unless some alkali be present; and the activity of an alkaline juice is arrested by acidification, and hindered by neutralization."[1] Therefore it seems highly probable that the innumerable calciferous cells, which are poured from the four posterior glands into the alimentary canal of worms, serve to neutralise more or less completely the acids there generated by the half-decayed leaves. We have seen that these cells are instantly dissolved by a small quantity of acetic acid, and as they do not always suffice to neutralise the contents of even the upper part of the alimentary canal, the lime is perhaps aggregated into concretions in the anterior pair of glands, in order that some may be

  1. M. Foster, Ibid. p. 200.