from the three responding pulvini. The electric connections are so made that the same tetanising shock is applied on the three petioles, very young, fully grown, and very old. The secondary coil is gradually pushed in till the leaves exhibit responsive fall. The fully grown leaf was the first to respond, the velocity of transmission being 23 mm. per second. The secondary coil had to be pushed nearer the primary through 6 cm. before excitation could be effectively transmitted through the young petiole; for the oldest leaf still stronger stimulus was necessary, since in this case the secondary had to be pushed through an additional distance of 4 cm. for effective transmission of excitation. I also determined the relative values of the minimal intensity of stimulus, effective in causing transmission of excitation in the three cases. Adopting as before the intensity of electric stimulus which causes bare perception in a human being as the unit, I find that the effective stimulus for a fully grown young petiole is 0.3 unit, while the very young required 2.5 units, and the very old 5 units. Hence it may be said that the conducting power of a very young is an eighth, and of the very old one-sixteenth of the conductivity of the fully grown young specimen.
It will thus be seen that the conducting power of a very young petiole is feebler than in a fully grown specimen. The conducting tissue, it is true, is present, but the power of conduction has not become fully developed. This power is, as we shall see later, conferred by the stimulus of the environment. In a very old specimen the diminution of conducting power is due to the general physiological decline.
EFFECT OF DESICCATION ON CONDUCTING TISSUES.
I have already shown that transmission in the plant is a process fundamentally similar to that taking place in
